Distribution
Austria, Ukraine

Native to: Afghanistan, Albania, Algeria, Altay, Assam, Austria, Baleares, Baltic States, Belarus, Belgium, Bulgaria, Buryatiya, Central European Rus, China North-Central, China South-Central, China Southeast, Chita, Corse, Cyprus, Czechoslovakia, Denmark, East European Russia, Finland, France, Germany, Great Britain, Greece, Hungary, Inner Mongolia, Iran, Ireland, Irkutsk, Italy, Japan, Kazakhstan, Khabarovsk, Kirgizstan, Korea, Krasnoyarsk, Krym, Lebanon-Syria, Manchuria, Morocco, Nepal, Netherlands, North Caucasus, North European Russi, Northwest European R, Norway, Pakistan, Poland, Portugal, Primorye, Romania, Sakhalin, Sardegna, Sicilia, Spain, Sweden, Switzerland, Tadzhikistan, Taiwan, Tibet, Transcaucasus, Turkey, Turkey-in-Europe, Tuva, Ukraine, West Himalaya, Xinjiang, Yakutskiya, Yugoslavia.

Roots not proliferous. Stems short-creeping to erect, often branched; scales very dark brown throughout, narrowly deltate, 1-3 × 0.1-0.25 mm, margins with widely spaced teeth. Leaves monomorphic. Petiole reddish brown proximally, green distally, dull, 1-9 cm, (1/2-)1-2 times length of blade; indument of dark brown, narrowly deltate scales proximally grading into multicellular hairs. Blade deltate-ovate to obovate or oblanceolate, 1-2(-3)-pinnate to 2-pinnate-pinnatifid, 2-6 × 1-4 cm, somewhat thick, glabrous; base obtuse; apex acute to rounded, not rooting. Rachis green, dull, glabrous except for very sparse, minute hairs. Pinnae in 2-4 pairs, deltate-ovate to obdeltate; proximal (largest) pinnae 7-30 × 5-20 mm; base truncate to acute; margins finely (sometimes coarsely) incised; apex rounded to acute. Veins free, evident. Sori as many as 30 or more per pinna, usually 1-5 per segment, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2 n = 144.
Limestone (or calcareous shale) cliffs and boulders, rarely invading masonry; 0-1000 m.